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EXHIBIT 264

Makarkinia

Makarkinia

Prehistoric insect

Makarkinia was a large Cretaceous lacewing known from distinctive fossil wings, including ringed hindwing eyespots. [1]

Evidence profile
PERIODEarly Cretaceous[2]
DIETPlant-fluid feeding inferredEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDBrazil[2]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Makarkinia

Makarkinia was an extinct lacewing, rather than a butterfly or dinosaur. A 2016 study described a hindwing of M. kerneri with dense veins and a dark eyespot surrounded by three closely spaced rings. The preserved wing measures 78 millimeters, while the researchers estimated a complete length of 100 to 120 millimeters. Those numbers describe one wing; they must not be presented as the insect’s body length or total wingspan. [D1]

A Brazilian fossil catalog lists three species within the genus: M. adamsi, M. irmae and M. kerneri. All come from the Aptian Crato Formation. The older name Panfilovia adamsi refers to the species later placed in Makarkinia. That change does not make every species of Panfilovia the same animal. Careful attention to names prevents a renamed fossil from being counted as a second exhibit. [D2]

A later research abstract reconsidered relationships among these remarkable lacewings. It described Makarkinia as probably belonging in the subfamily Kalligrammatinae. The qualifying word shows that part of its placement remains uncertain. Its general relationship to other large lacewings is useful, but cannot supply missing body parts as directly preserved evidence. Reconstructing a complete living insect requires comparative choices that should remain separate from the measured fossil wing. [D3]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Its preserved wing is measurable; total body length, exact wingspan and life history remain uncertain. [D2]

Profile scope: Genus — Makarkinia; gallery pattern anchored to the M. kerneri hindwing holotype SMNS70287. Complete body anatomy is restored comparatively. [D1]

Time
Early Cretaceous
Fossil locations in this exhibit
Brazil
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Size reference

Source supported

78 mm preserved hindwing; 100–120 mm estimated complete wing

M. kerneri holotype reference. Neither measurement is body length or wingspan. [D1]

Body mass

Not established

Not established here

The reviewed wing description gives no body-mass estimate. [D1]

Diet

Inferred

Plant-fluid feeding inferred

This is a kalligrammatid comparison; M. kerneri preserves no mouthparts or identified meal. [D1]

Habitat

Inferred

Terrestrial Crato landscape inferred

The Aptian lake-burial horizon does not establish an exact living perch. [D2]

Movement

Inferred

Flight inferred

The described adult wing supports flight; detailed leg movement and speeds are unknown. [D1]

Distinctive anatomy

Source supported

Dense wing veins and ringed eyespot

M. kerneri has three closely spaced dark rings around its hindwing eyespot; the body is unpreserved. [D1]

Reproduction & life together

Reproduction

Not established

Not established here

The reviewed adult-wing material does not identify eggs, larvae or nests. [D2]

Courtship

Not established

Not established here

The catalog’s taxonomic records do not demonstrate courtship displays. [D2]

Social behavior

Not established

Not established here

Distinct fossil species do not establish swarms, cooperation or colonies. [D2]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. A new gigantic lacewing species (Insecta: Neuroptera) from the Lower Cretaceous of Brazil confirms the occurrence of Kalligrammatidae in the AmericasCretaceous Research; Bechly & Makarkin, 2016 · Accessed 2026-10-09Full original author-institution PDF inspected, including SMNS70287 diagnosis, preserved/estimated wing lengths, eyespot rings, Panfilovia adamsi recombination and figures 2–3 actual wing/venation. · Inspected: page
  2. An annotated catalog of the fossil Neuropterida (Insecta: Raphidioptera, Megaloptera, Neuroptera) of BrazilRevista Brasileira de Entomologia; Martins et al., 2022 · Accessed 2026-10-09Full original catalog PDF inspected; Makarkinia treatment lists adamsi, irmae and kerneri, species former combination, Crato Aptian horizons and type repositories. Known fossil stage is distinguished from unreported life-history evidence. · Inspected: page
  3. Redescription of Kalligramma haeckeli Walther, 1904 allows the redefinition of KalligrammatinaeZootaxa; Makarkin & Ansorge, 2025 · Accessed 2026-10-09Original publisher abstract inspected. Probably Makarkinia wording retained; genus is not among the proposed Kalligramma genus synonyms. Existing catalog prose on this URL is included in the combined budget. · Inspected: abstract

THE STORY IN THE FOSSILS

Placement remains qualified

The 2025 abstract places Makarkinia probably within Kalligrammatinae, retaining uncertainty in its subfamily assignment. [3]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. A new gigantic lacewing species (Insecta: Neuroptera) from the Lower Cretaceous of Brazil confirms the occurrence of Kalligrammatidae in the AmericasCretaceous Research; Bechly & Makarkin, 2016 · Accessed 2026-10-09Full original author-institution PDF inspected, including SMNS70287 diagnosis, preserved/estimated wing lengths, eyespot rings, Panfilovia adamsi recombination and figures 2–3 actual wing/venation. · Inspected: page
  2. An annotated catalog of the fossil Neuropterida (Insecta: Raphidioptera, Megaloptera, Neuroptera) of BrazilRevista Brasileira de Entomologia; Martins et al., 2022 · Accessed 2026-10-09Full original catalog PDF inspected; Makarkinia treatment lists adamsi, irmae and kerneri, species former combination, Crato Aptian horizons and type repositories. Known fossil stage is distinguished from unreported life-history evidence. · Inspected: page
  3. Redescription of Kalligramma haeckeli Walther, 1904 allows the redefinition of KalligrammatinaeZootaxa; Makarkin & Ansorge, 2025 · Accessed 2026-10-09Original publisher abstract inspected. Probably Makarkinia wording retained; genus is not among the proposed Kalligramma genus synonyms. Existing catalog prose on this URL is included in the combined budget. · Inspected: abstract

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