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EXHIBIT 219

Yi

Yi qi

Dinosaur

Yi was a small feathered dinosaur with wrist-supported wing membranes, a different construction from a bird’s feathered wings. [1]

Evidence profile
PERIODLate Jurassic[1]
DIETOmnivoreEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDChina[1]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Yi

Yi was a small dinosaur from Jurassic China with an unusual combination of feathers and wing membranes. It belonged to the scansoriopterygids, a group of small theropods. Rod-like structures extended from its wrists beside the long fingers. These supports make its wing construction different from a bird’s feathered flight surfaces. The fossil is incomplete, so artists must reconstruct some missing body parts and membrane boundaries. [D1]

The type specimen is estimated to have weighed about three hundred and eighty grams. Researchers calculated that value using an equation; they could not put the living dinosaur on a scale. Closed joints between parts of its vertebrae suggest that it was an adult. The fossil also preserves stiff, simple feathers and sheet-like tissue, which provide clues to the covering and unusual surfaces of its body. [D2]

Later researchers tested possible wings in aerodynamic models. The results suggest that Yi was unlikely to fly by powering strong wingbeats. Gliding was possible but limited, and the outcome depended on the reconstructed wing shape. A wooded setting offers possible launch points and short gaps to cross. These calculations help test a way of moving, while leaving the animal’s exact movements and preferred habitat uncertain. [D3]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Closed vertebral sutures suggest adulthood in the type; its approximately 380 g mass is an empirical estimate. [D2]

Profile scope: Genus — Yi qi; partial skeleton STM31-2 from the Middle–Upper Jurassic Tiaojishan Formation of Hebei, China. [D1]

Time
Late Jurassic
Fossil locations in this exhibit
China
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Body length

Not established

Not established here

The inspected partial skeleton account does not establish a comparable whole-body length or species range. [D2]

Body mass

Inferred

About 380 g (type-individual model)

An empirical equation estimates STM31-2; vertebral fusion suggests adulthood. This is not a weighed animal or species range. [D2]

Diet and feeding

Not established

Not established here

The catalog’s omnivore grouping is provisional. The reviewed fossil account does not demonstrate a specific meal. [D1]

Habitat and setting

Inferred

Northern Chinese Jurassic forest setting

Forest-gap gliding is a proposed ecological scenario; burial setting does not fix the precise canopy habitat. [D3]

Movement

Inferred

Limited gliding modeled

Powered flight is unlikely in the tested models; exact wing outline affects calculated performance. [D3]

Recognizable anatomy

Source supported

Wrist rods and wing membranes

Filamentous feathers accompany sheet-like tissue; reconstructed complete membrane boundaries remain uncertain. [D2]

Reproduction & life together

Reproduction and nesting

Not established

Not established here

The reviewed partial skeleton and focused searches do not identify assigned eggs, nests or parental care. [D1]

Mating and courtship

Not established

Not established here

No reviewed fossil establishes display gestures, mating posture or sex-specific coloration. [D1]

Social behavior

Not established

Not established here

The described specimen does not demonstrate a flock, communal roost or cooperative behavior. [D1]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. A bizarre Jurassic maniraptoran theropod with preserved evidence of membranous wings (Xu et al., 2015)Nature 521:70–73 · Accessed 2026-10-05Original publisher abstract and extended fossil figures/captions inspected. Yi qi STM31-2 is a distinctive scansoriopterygid partial skeleton, with unusual styliform wrist rods, filamentous feathers and membrane-like soft tissue. Complete wing outlines and missing body regions require inference. Subscription main osteological text was not represented as fully inspected. · Inspected: excerpt
  2. New scansoriopterygid theropod Yi qi — BCAS research report (2015)Bulletin of the Chinese Academy of Sciences 29(2) · Accessed 2026-10-05Indexed institutional PDF excerpt inspected: type adult status inferred from closed neurocentral sutures; body mass approximately 380 g via an empirical equation; unusual wrist rods, filaments and sheet-like soft tissue described. Complete tail/length and genus-specific reproduction are not supplied. · Inspected: excerpt
  3. Aerodynamics Show Membrane-Winged Theropods Were a Poor Gliding Dead-end (Dececchi et al., 2020)iScience 23:101574 · Accessed 2026-10-05Full original article inspected through official NCBI BioC XML PMC7756141. Laser-stimulated fluorescence and parameter-sensitive flight models test Yi and Ambopteryx: powered flight unlikely, gliding limited. Arboreal/forest-gap scenarios and diet are hypotheses, not observed behavior. Both exhibit citations share one source word budget. · Inspected: page

THE STORY IN THE FOSSILS

Evidence behind Yi

Its wings combined membranes and unusual wrist supports. Missing tissue still makes the complete wing boundary a reconstruction. [2]

The small mass estimate belongs to the type specimen, whose adulthood is inferred from fused vertebral joints. [2]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. A bizarre Jurassic maniraptoran theropod with preserved evidence of membranous wings (Xu et al., 2015)Nature 521:70–73 · Accessed 2026-10-05Original publisher abstract and extended fossil figures/captions inspected. Yi qi STM31-2 is a distinctive scansoriopterygid partial skeleton, with unusual styliform wrist rods, filamentous feathers and membrane-like soft tissue. Complete wing outlines and missing body regions require inference. Subscription main osteological text was not represented as fully inspected. · Inspected: excerpt
  2. New scansoriopterygid theropod Yi qi — BCAS research report (2015)Bulletin of the Chinese Academy of Sciences 29(2) · Accessed 2026-10-05Indexed institutional PDF excerpt inspected: type adult status inferred from closed neurocentral sutures; body mass approximately 380 g via an empirical equation; unusual wrist rods, filaments and sheet-like soft tissue described. Complete tail/length and genus-specific reproduction are not supplied. · Inspected: excerpt
  3. Aerodynamics Show Membrane-Winged Theropods Were a Poor Gliding Dead-end (Dececchi et al., 2020)iScience 23:101574 · Accessed 2026-10-05Full original article inspected through official NCBI BioC XML PMC7756141. Laser-stimulated fluorescence and parameter-sensitive flight models test Yi and Ambopteryx: powered flight unlikely, gliding limited. Arboreal/forest-gap scenarios and diet are hypotheses, not observed behavior. Both exhibit citations share one source word budget. · Inspected: page

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