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EXHIBIT 221

Archaboilus

Archaboilus

Prehistoric insect

Archaboilus was a Jurassic katydid relative whose fossil wings preserve clues to ancient insect calls. [3]

Evidence profile
PERIODEarly–Middle Jurassic[4]
DIETNot establishedEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDKyrgyzstan, Russia, China[4]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Archaboilus

Archaboilus was an insect relative of katydids, living long before modern gardens and lawns. A species called Archaboilus musicus came from Middle Jurassic rocks at Daohugou in China. Its best-known fossil is an incomplete pair of male forewings. Researchers estimated a forewing length of about 72 millimeters. That measures one reconstructed wing, not the length of the whole insect. [D1]

Tiny features on those wings helped researchers work out how the animal might have made a call. A row of teeth could rub against a structure on the other wing. Comparisons with living insects produced a model of a musical tone. The researchers interpreted singing as a way to attract a mate, but no fossil records the insect actually singing or meeting a partner. [D3]

A2026 study compared several Jurassic insects and found more than one kind of sound-making arrangement. Another species referred to Archaboilus, A. polyneurus, had tooth-spacing patterns consistent with changing its call. A. musicus remains a low-frequency example; findings about ultrasonic calls in other insects do not turn every member of this genus into an ultrasonic singer. Fossil anatomy supplies clues, while the sound itself remains a reconstruction. [D2]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Sound estimates come from wing mechanics and living comparisons; no Jurassic sound recording survives. [D2]

Profile scope: Genus — Archaboilus; gallery outline anchored to A. musicus, an incomplete male forewing pair from Daohugou, China. [D1]

Time
Early–Middle Jurassic
Fossil locations in this exhibit
Kyrgyzstan, Russia, China
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Size reference

Source supported

About72 mm estimated forewing

Estimate for the incomplete A. musicus male type; not body length, wingspan or a genus range. [D1]

Body mass

Not established

Not established here

Reviewed wing-based taxonomy provides no fossil-individual body-mass estimate. [D4]

Diet

Not established

Not established here

The focal wing pair preserves no mouthparts or meal establishing diet. [D1]

Habitat

Inferred

Jurassic woodland inferred

Forest and possible nocturnal activity are acoustic comparisons; burial does not identify its exact perch. [D3]

Movement

Inferred

Flight inferred

Forewings indicate a winged insect; flight speed and jumping performance remain unmeasured. [D3]

Distinctive anatomy

Source supported

Veined sound-making forewings

A toothed file supports stridulation; the focal type lacks the complete head, legs and body. [D1]

Reproduction & life together

Reproduction

Not established

Not established here

The reviewed adult-wing descriptions identify no attributable eggs or juvenile series. [D4]

Courtship

Inferred

Mate-attraction call inferred

The2012 model estimates a 6.4 kHz tone for A. musicus; comparison suggests mate attraction, not a preserved mating sequence. [D3]

Social behavior

Not established

Not established here

Modeled species calls do not demonstrate a cooperative group or an observed chorus. [D2]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. Wing stridulation in a Jurassic katydid (Insecta, Orthoptera) produced low-pitched musical calls to attract femalesPNAS; Gu et al., 2012 · Accessed 2026-10-06Original author-hosted article and supplement inspected: A. musicus type, incomplete72 mm estimated forewing, stridulatory file, modeled6.4 kHz call and restoration limits. · Inspected: page
  2. Reconstruction of an extinct soundscape reveals ultrasonic communication in the JurassicPNAS; Gu et al., 2026 · Accessed 2026-10-06Original full article captured via EuropePMC and relevant Results/Discussion inspected. A. musicus low-frequency comparison and A. polyneurus modulation are kept distinct; ultrasonic findings from other species are not transferred. · Inspected: page
  3. Fossil cricket reveals Jurassic love songUniversity of Bristol, 2012 · Accessed 2026-10-06Full institutional research-team release inspected: mechanical reconstruction, mating-call interpretation, likely forest/nocturnal ecology. Suggested named predators and confident reconstructed timing are not adopted as direct observations. · Inspected: page
  4. Systematics Review and Phylogeny of Cyrtophyllitinae Zeuner, 1935 sensu Gorochov, Jarzembowski & Coram, 2006 (Ensifera, Haglidae), with Description of Two New SpeciesInsects; Gu, Ren & Chen, 2024 · Accessed 2026-10-06Full original taxonomy sections, Table1 and revised diagnosis inspected: Archaboilus is Haglidae/Archaboilinae, Early–Middle Jurassic Kyrgyzstan, Russia, China; A. polyneurus placement retained as probable. · Inspected: page

THE STORY IN THE FOSSILS

A call hidden in a wing

Different tooth spacing can suggest different sound patterns. The2026 study treated A. polyneurus separately from the familiar A. musicus reconstruction, rather than assigning one pitch to every Archaboilus species. [2]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. Wing stridulation in a Jurassic katydid (Insecta, Orthoptera) produced low-pitched musical calls to attract femalesPNAS; Gu et al., 2012 · Accessed 2026-10-06Original author-hosted article and supplement inspected: A. musicus type, incomplete72 mm estimated forewing, stridulatory file, modeled6.4 kHz call and restoration limits. · Inspected: page
  2. Reconstruction of an extinct soundscape reveals ultrasonic communication in the JurassicPNAS; Gu et al., 2026 · Accessed 2026-10-06Original full article captured via EuropePMC and relevant Results/Discussion inspected. A. musicus low-frequency comparison and A. polyneurus modulation are kept distinct; ultrasonic findings from other species are not transferred. · Inspected: page
  3. Fossil cricket reveals Jurassic love songUniversity of Bristol, 2012 · Accessed 2026-10-06Full institutional research-team release inspected: mechanical reconstruction, mating-call interpretation, likely forest/nocturnal ecology. Suggested named predators and confident reconstructed timing are not adopted as direct observations. · Inspected: page
  4. Systematics Review and Phylogeny of Cyrtophyllitinae Zeuner, 1935 sensu Gorochov, Jarzembowski & Coram, 2006 (Ensifera, Haglidae), with Description of Two New SpeciesInsects; Gu, Ren & Chen, 2024 · Accessed 2026-10-06Full original taxonomy sections, Table1 and revised diagnosis inspected: Archaboilus is Haglidae/Archaboilinae, Early–Middle Jurassic Kyrgyzstan, Russia, China; A. polyneurus placement retained as probable. · Inspected: page

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