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EXHIBIT 29

Citipati

Citipati

A dinosaur preserved with its arms spread over a nest.

Evidence profile
PERIODLate Cretaceous[1]
DIETOmnivoreInferred from related anatomy [1]
TOTAL LENGTHNot established hereComparable estimate unavailable
FOSSILS FOUNDMongolia[1]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Citipati

Citipati was a two-legged dinosaur from Late Cretaceous Mongolia with a beak rather than teeth. The Natural History Museum lists an omnivorous diet, meaning that both plant and animal foods are proposed. That broad interpretation is useful for imagining its place in an ecosystem, but it is not a list of meals directly recovered from every Citipati fossil. [D1]

One specimen nicknamed Big Mama gives unusually strong evidence about nesting. Its skeleton was preserved curled over a nest of eggs, a position interpreted as brooding. Brooding means sitting over eggs to help incubate or protect them. The roughly 75-million-year-old Mongolian fossil was announced in 1995 and later identified as Citipati when that genus was named in 2001. [S1]

This is much more informative than finding scattered eggs alone: the adult's position connects an animal with a particular nest. Even so, a single frozen moment cannot reveal its whole breeding season. The fossil supports care at the egg stage, while leaving questions about later chick care, courtship and division of parental duties unanswered in this account. [S1]

Overview sources
  1. Were dinosaurs good parents?Natural History Museum, London · Accessed 2026-10-04Inspected: page

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Citipati is an oviraptorid best known for fossils positioned over egg clutches in a bird-like brooding posture. Its well-preserved skull also documents extensive internal air spaces, while its precise diet and mating system remain less certain.

Profile scope: Genus: Citipati; brooding and skull evidence concerns C. osmolskae.

Time
Late Cretaceous
Fossil locations in this exhibit
Mongolia
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Body length

Inferred

About 2.1 m; museum reference

NHM gives a rounded genus figure with no specified specimen or life stage. It is not a maximum size claim. [D1]

Body mass

Not established

Not established here

The reviewed sources do not establish a numerical body-mass estimate for this record.

Diet

Inferred

Possible omnivore; exact foods unresolved

NHM lists omnivory, but the reviewed material does not establish a menu through gut contents. Beak form alone does not reveal every food consumed. [D1]

Habitat & fossil setting

Source supported

Djadokhta Formation, Ukhaa Tolgod, Mongolia

DigiMorph identifies the formation and locality for the C. osmolskae holotype. This is fossil context rather than a demonstrated habitat preference. [D3]

Movement

Inferred

Bipedal theropod; gait unreviewed

Museum skeletal reconstructions show a two-legged theropod. The inspected sources do not establish a running speed or complete gait model. [D2]

Distinctive anatomy

Source supported

Beaked skull with extensive pneumatic recesses

CT work on the holotype documents numerous skull recesses interpreted as air spaces. The skull differs from the true Oviraptor holotype. [D3]

Reproduction & life together

Reproduction & nesting

Source supported

Egg laying; bird-like brooding posture

C. osmolskae fossils are positioned over nests with the forearms spread around the eggs. Which sex brooded and how parental duties were divided remain unresolved. [D2]

Courtship & mating

Not established

Not established here

The reviewed sources do not establish a mating ritual, pair bond or breeding season for this taxon.

Social behavior

Not established

Not established here

The reviewed sources do not establish a taxon-specific group structure or social hierarchy.

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. Citipati — Dino DirectoryNatural History Museum, London · Accessed 2026-10-02Museum genus reference; life stage and specimen not specified. · Inspected: page
  2. Get to Know a Dino: Citipati osmolskaeAmerican Museum of Natural History · Accessed 2026-10-02C. osmolskae brooding fossil and limits of parental-sex interpretations. · Inspected: page
  3. Citipati osmolskae — expert CT annotationUniversity of Texas DigiMorph; Clark, Norell & Rowe (2002) · Accessed 2026-10-02C. osmolskae holotype skull; comparison with O. philoceratops and rejected older referral. · Inspected: page

THE STORY IN THE FOSSILS

Meet Citipati.

A Citipati skeleton was preserved over the centre of a nest with its forearms extended around the eggs. The posture resembles brooding in living birds. It provides unusually direct evidence that nesting behavior was already present among dinosaurs outside the modern bird lineage. [2]

Museum and Mongolian Academy of Sciences researchers discovered the first Citipati specimens at Ukhaa Tolgod in 1993. Its well-preserved skull also reveals extensive internal air spaces. Comparisons with that skull helped distinguish true Oviraptor from a different, frequently illustrated oviraptorid skeleton. [2] [3]

A note on scale and time

We have not established a comparable adult total-length estimate for this taxon in this draft. Specimen measurements in the text describe those individuals. The cited dates are approximately 81–75 million years ago. Approximate museum catalogue reference interval for the genus, not an exact duration for every species. Unknown does not mean zero.

Sources & editorial notes

Sources and evidence notes for this exhibit.

  1. Citipati — Dino DirectoryNatural History Museum, London · Accessed 2026-10-02Profile and Taxonomic details. Genus diet is an educational interpretation; 2.1 m is not given an adult life-stage scope.
  2. Get to Know a Dino: Citipati osmolskaeAmerican Museum of Natural History · Accessed 2026-10-02Protective Pose; Doting Dads?; In Good Company. Fossil pose supports nesting behavior; parental sex and division of duties are explicitly unknown. Feathered model is a reconstruction.
  3. Citipati osmolskae — expert CT annotation from Clark, Norell and Rowe (2002)DigiMorph, University of Texas at Austin · Accessed 2026-10-02Expert annotation, paragraphs quoted from the original cranial paper, especially poorly preserved Oviraptor holotype, elongated rostrum/dentary and reassignment of IGM 100/42. Skull record is species-specific.

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