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EXHIBIT 111

Dacentrurus

Dacentrurus armatus

Dacentrurus was a heavily built European stegosaur, combining bony armor with a substantial pelvis and limbs. [2]

Evidence profile
PERIODLate Jurassic[2]
DIETHerbivoreEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDEngland, UK, Spain[6]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Dacentrurus

Dacentrurus was a plant-eating stegosaur from Late Jurassic Europe. Its museum reference of about six metres gives a general sense of scale without naming a measured complete adult. The genus has a long research history, and its profile depends on carefully deciding which later discoveries can be compared with its original fossil material. [D1]

A proposed merger with the long-necked Miragaia remains contested. Research arguing that Miragaia is distinct identifies problems with that suggested synonymy. A synonym is another name interpreted as referring to the same animal. Because the disagreement is unresolved, Miragaia's distinctive neck and life-history claims should not automatically be transferred to Dacentrurus. [D4]

Spanish trackways called Deltapodus suggest that some stegosaurs traveled together. Tracks preserve movement more directly than a mounted skeleton, but identifying the exact genus that made them is difficult. They do not securely demonstrate Dacentrurus herds or one measured walking speed. Keeping the trackmaker uncertain lets the footprints inform stegosaur behavior without giving a named animal more detailed habits than the evidence supports. [D8]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Its incomplete skeletons leave the full armor arrangement uncertain. Recent cranial discoveries expand its anatomy, while disagreements over related genera still affect its reconstruction. [D3]

Profile scope: D. armatus, anchored by the Swindon type and recent Spanish referrals; disputed Miragaia material is kept separate. [D3] [D4]

Time
Late Jurassic
Fossil locations in this exhibit
England, UK, Spain
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Length

Inferred

About 6 m (museum reference)

The museum gives a genus-level educational reconstruction, without a named individual, method or maturity assessment. It is not a measured adult range. [D1]

Mass

Inferred

About 7.4 tonnes (limb model)

Benson models type BMNH46013; its crushed femur required an estimated circumference. This uncertain individual reconstruction is not an adult population average. [D7]

Diet and feeding

Source supported

Plant-eater

The museum identifies herbivory; particular food plants and a genus-specific feeding height are not established here. [D1]

Habitat and range

Source supported

Late Jurassic European land environments

Spanish referrals occur in Villar del Arzobispo coastal river deposits, including wetlands and floodplains. Burial environments do not establish an exclusive habitat preference. [D6]

Movement

Inferred

Four-legged walking inferred

Its stegosaur limb plan supports quadrupedality. Trackways assigned only to stegosaurs cannot identify a Dacentrurus speed or gait directly. [D2]

Distinctive anatomy

Source supported

Substantial pelvis and limbs; plates and spines

The type preserves a large pelvis and limb material; newly referred Spanish cranial remains add skull information. A complete, securely arranged armor series is lacking. [D2] [D3]

Reproduction & life together

Reproduction and care

Not established

Not established here

Focused review found no securely assigned eggs, nests, incubation pattern or parental care for D. armatus. Miragaia growth results are not transferred to it.

Courtship and mating

Not established

Not established here

Plates and spines alone do not demonstrate sexual dimorphism, mate choice or a particular mating ritual. No such genus-specific evidence was established in this review.

Social behavior

Inferred

Possible group travel; trackmaker uncertain

Spanish Deltapodus trackways suggest stegosaur grouping. A 2026 study proposes D. armatus as the likely maker; neither genus identity nor lasting herds are demonstrated. [D8] [D6]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. DacentrurusNatural History Museum, London · Accessed 2026-10-02Museum profile inspected: educational 6 m length, herbivory and Late Jurassic European occurrence. Individual, age and length method are not specified. · Inspected: page
  2. Stegosauria: a historical review of the body fossil record and phylogenetic relationshipsSwiss Journal of Geosciences; Maidment (2010) · Accessed 2026-10-02Original 12-page review inspected, including the Swindon type, preoccupied Omosaurus name, stegosaur anatomy and quadrupedality. Historical referrals are checked against newer studies. · Inspected: page
  3. New insights into the phylogeny and skull evolution of stegosaurian dinosaurs: An extraordinary cranium from the European Late Jurassic (Dinosauria: Stegosauria)Vertebrate Zoology; Sánchez-Fenollosa and Cobos (2025) · Accessed 2026-10-02Published full text inspected through a publisher/author-uploaded copy. Spanish cranium MAP-9029, D. armatus referrals and synonymy including Miragaia; latter interpretation is disputed by Costa and colleagues. · Inspected: page
  4. New evidence from Miragaia longicollum MG 4863Comunicações Geológicas; Costa, Maidment, Sequero and Crespo (2025) · Accessed 2026-10-02Published full PDF inspected via a copy of the institutional article; anatomical comparisons support Miragaia as distinct and identify problems with the proposed synonymy. Later portal access was bot-blocked. · Inspected: page
  5. Dacentrurine Stegosaurs in North America: New Occurrences from the Upper Jurassic of USA (Morrison Formation)Diversity; Costa (2026) · Accessed 2026-10-02Published 26-page full PDF inspected. Retains Miragaia, including M. longispinus, as distinct; North American referrals do not independently establish a Dacentrurus range there. · Inspected: page
  6. The plated dinosaurs (Ornithischia, Stegosauria) of eastern Iberia (Spain): Taxonomy, diversity, and ecologyPalaeontologia Electronica; Sánchez-Fenollosa, Cobos and Suñer (2026) · Accessed 2026-10-02Published 44-page full PDF inspected, especially taxonomy and ecology. D. armatus referrals, Villar del Arzobispo depositional environments and proposed Deltapodus trackmaker identity; this authorship continues the disputed Miragaia synonymy. · Inspected: page
  7. Rates of Dinosaur Body Mass Evolution Indicate 170 Million Years of Sustained Ecological Innovation on the Avian Stem LineagePLOS Biology; Benson et al. (2014) · Accessed 2026-10-02Methods and Dataset S1 XLS inspected. Dacentrurus armatus BMNH46013 is modeled at 7,374.39 kg; femoral circumference is estimated because the type femur is crushed, rather than directly measured. · Inspected: page
  8. The complexity of tracking stegosaurs and their gregarious behaviorScientific Reports; Castanera, Mampel and Cobos (2024) · Accessed 2026-10-02Primary article text and discussion inspected through indexed publisher/PMC text; institutional abstract also inspected. Deltapodus trackway orientation and possible stegosaur grouping do not prove a genus-level trackmaker. Institutional PDF download timed out. · Inspected: excerpt

THE STORY IN THE FOSSILS

Meet Dacentrurus

The Swindon fossil was originally named Omosaurus armatus. Because Omosaurus was already occupied, the animal acquired the name Dacentrurus. Its incomplete type remains the anchor for identifying later discoveries. [2]

Spanish finds now supply cranial evidence and several additional referrals. The proposed merger with long-necked Miragaia remains contested, so those fossils do not automatically establish the neck or life history of D. armatus. [3] [4] [5]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. DacentrurusNatural History Museum, London · Accessed 2026-10-02Museum profile inspected: educational 6 m length, herbivory and Late Jurassic European occurrence. Individual, age and length method are not specified. · Inspected: page
  2. Stegosauria: a historical review of the body fossil record and phylogenetic relationshipsSwiss Journal of Geosciences; Maidment (2010) · Accessed 2026-10-02Original 12-page review inspected, including the Swindon type, preoccupied Omosaurus name, stegosaur anatomy and quadrupedality. Historical referrals are checked against newer studies. · Inspected: page
  3. New insights into the phylogeny and skull evolution of stegosaurian dinosaurs: An extraordinary cranium from the European Late Jurassic (Dinosauria: Stegosauria)Vertebrate Zoology; Sánchez-Fenollosa and Cobos (2025) · Accessed 2026-10-02Published full text inspected through a publisher/author-uploaded copy. Spanish cranium MAP-9029, D. armatus referrals and synonymy including Miragaia; latter interpretation is disputed by Costa and colleagues. · Inspected: page
  4. New evidence from Miragaia longicollum MG 4863Comunicações Geológicas; Costa, Maidment, Sequero and Crespo (2025) · Accessed 2026-10-02Published full PDF inspected via a copy of the institutional article; anatomical comparisons support Miragaia as distinct and identify problems with the proposed synonymy. Later portal access was bot-blocked. · Inspected: page
  5. Dacentrurine Stegosaurs in North America: New Occurrences from the Upper Jurassic of USA (Morrison Formation)Diversity; Costa (2026) · Accessed 2026-10-02Published 26-page full PDF inspected. Retains Miragaia, including M. longispinus, as distinct; North American referrals do not independently establish a Dacentrurus range there. · Inspected: page
  6. The plated dinosaurs (Ornithischia, Stegosauria) of eastern Iberia (Spain): Taxonomy, diversity, and ecologyPalaeontologia Electronica; Sánchez-Fenollosa, Cobos and Suñer (2026) · Accessed 2026-10-02Published 44-page full PDF inspected, especially taxonomy and ecology. D. armatus referrals, Villar del Arzobispo depositional environments and proposed Deltapodus trackmaker identity; this authorship continues the disputed Miragaia synonymy. · Inspected: page
  7. Rates of Dinosaur Body Mass Evolution Indicate 170 Million Years of Sustained Ecological Innovation on the Avian Stem LineagePLOS Biology; Benson et al. (2014) · Accessed 2026-10-02Methods and Dataset S1 XLS inspected. Dacentrurus armatus BMNH46013 is modeled at 7,374.39 kg; femoral circumference is estimated because the type femur is crushed, rather than directly measured. · Inspected: page
  8. The complexity of tracking stegosaurs and their gregarious behaviorScientific Reports; Castanera, Mampel and Cobos (2024) · Accessed 2026-10-02Primary article text and discussion inspected through indexed publisher/PMC text; institutional abstract also inspected. Deltapodus trackway orientation and possible stegosaur grouping do not prove a genus-level trackmaker. Institutional PDF download timed out. · Inspected: excerpt

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