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EXHIBIT 54

Dreadnoughtus

Dreadnoughtus schrani

An exceptionally complete giant that was still growing.

Evidence profile
PERIODLate Cretaceous[1]
DIETHerbivoreInferred from related anatomy [1]
TOTAL LENGTHNot established hereComparable estimate unavailable
FOSSILS FOUNDSouthern Patagonia, Argentina[1]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Dreadnoughtus

Dreadnoughtus schrani was an enormous long-necked titanosaur from Late Cretaceous Patagonia, in southern Argentina. Two individuals preserve complementary parts of its skeleton. Their bones include substantial portions of the limbs, shoulder region, pelvis and tail, giving researchers unusually useful evidence for reconstructing a giant sauropod. The remains are still incomplete, particularly where a complete head and feet would be needed. [S1]

Its weight is especially sensitive to the method used. One estimate scaled up from limb measurements; another built a three-dimensional body around the skeleton and allowed different amounts of soft tissue. The latter produced considerably lower estimates. Neither weighs the living dinosaur directly. Their disagreement shows why a giant’s famous weight should be reported with its reconstruction method, rather than treated as a precise measurement. [S2]

Microscopic bone structure indicates that the larger described individual was still growing when it died. It therefore cannot automatically define the maximum adult size of the species. The bones also preserve muscle attachment details that help researchers understand how its body was supported. Finding two animals provides anatomical comparisons, but does not by itself establish a herd or explain their relationships during life. [S1]

Overview sources
  1. A Gigantic, Exceptionally Complete Titanosaurian Sauropod Dinosaur from Southern Patagonia, ArgentinaScientific Reports; Lacovara and colleagues (2014) · Accessed 2026-10-04Inspected: excerpt
  2. Scientists downsize the giant Dreadnoughtus dinosaurUniversity of Liverpool · Accessed 2026-10-04Inspected: excerpt

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Dreadnoughtus preserves unusually informative limb and trunk material for a giant titanosaur. Its holotype was still growing, and limb-based versus volumetric methods give sharply different masses for that same individual.

Profile scope: Species: Dreadnoughtus schrani; size estimates concern the growing holotype, not a fully grown adult range.

Time
Late Cretaceous
Fossil locations in this exhibit
Southern Patagonia, Argentina
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Length

Inferred

Holotype: about 26 m (reconstructed)

Drexel reports this reconstructed length for an individual whose bones indicate continued growth. It does not establish the size range of fully grown adults. [D1]

Mass

Debated

Holotype estimates differ: 22.1–38.2 versus 59.3 tonnes

The original limb-circumference method produced 59,300 kg; three volumetric models yielded roughly 22,100–38,200 kg. This disagreement concerns methods applied to one growing specimen, not an adult species range. [D1] [D2]

Diet and feeding

Inferred

Plant eater

The discovery team identifies a herbivorous titanosaur. Its proposed feeding envelope and food intake are reconstructions rather than preserved behavior. [D1]

Habitat and setting

Inferred

Southern Patagonian forest and floodplain reconstructed

Drexel describes a temperate forest setting and rapid burial after a river breached its levee. Depositional circumstances do not establish preferred habitat everywhere. [D1]

Movement

Inferred

Four-legged terrestrial reconstruction

Preserved fore- and hindlimb bones support quadrupedal reconstruction. A digital neutral pose is a modelling choice, not an observed walking cycle. [D1] [D2]

Distinctive anatomy

Source supported

Massive limbs with pronounced muscle attachments

The holotype retains many limb bones and clear attachment scars. Most of its skull and neck remain missing despite unusually good preservation for a giant titanosaur. [D1] [D2]

Reproduction & life together

Reproduction and care

Not established

Not established here

The reviewed sources do not establish nest structure, incubation or parental care for Dreadnoughtus schrani.

Courtship and mating

Not established

Not established here

The reviewed sources do not establish a courtship sequence or mating system for Dreadnoughtus schrani.

Social behavior

Not established

Not established here

Two individuals were recovered at the site. This association does not establish herd living, pair bonding or usual group size. [D1]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. Drexel Team Unveils Dreadnoughtus: A Gigantic, Exceptionally Complete Sauropod DinosaurDrexel University · Accessed 2026-10-02D. schrani discovery and holotype reconstruction; original mass claim subsequently challenged. · Inspected: page
  2. Downsizing a giant: re-evaluating Dreadnoughtus body massBiology Letters; Bates and colleagues (2015) · Accessed 2026-10-02Holotype volumetric modelling and comparison with the original limb-circumference estimate. · Inspected: excerpt

THE STORY IN THE FOSSILS

Meet Dreadnoughtus.

The Dreadnoughtus discovery recovered two skeletons in southern Patagonia. The larger individual preserved much of the skeleton outside the head, providing an unusually detailed view of a giant titanosaur. Drexel researchers estimated it at about 26 metres long and concluded that it was not yet fully grown. [1]

Its mass illustrates how different methods can disagree. The original limb-circumference calculation gave 59,300 kilograms, whereas a later volumetric study produced roughly 22,100–38,200 kilograms across its model variants. Those numbers concern a particular specimen and modelling choices, rather than a settled range for full-grown adults. [1] [2]

A note on scale and time

We have not established a comparable adult total-length estimate for this taxon in this draft. Specimen measurements in the text describe those individuals. The cited dates are approximately 77 million years ago. Approximate reference age, represented by equal endpoints; not a demonstrated full temporal range. Unknown does not mean zero.

Sources & editorial notes

Sources and evidence notes for this exhibit.

  1. Drexel Team Unveils Dreadnoughtus: A Gigantic, Exceptionally Complete Sauropod DinosaurDrexel University · Accessed 2026-10-024 September 2014 discovery release: D. schrani, southern Patagonia, approx. 77 Ma; titanosaur plant eater; 26 m and 59,300 kg refer to still-growing specimen; excavation and rapid burial interpretations. Claims of definitive record mass are not retained.
  2. Downsizing a giant: re-evaluating Dreadnoughtus body massBiology Letters / University of Brighton repository · Accessed 2026-10-02PDF pp. 1–4: Abstract, Methods, Results/Table 1, Discussion. Holotype model; missing cervical vertebrae, hand, skull and tail tip; volume/density models give 22,117–38,225 kg, challenging 59,300 kg limb-circumference estimate. These are method-dependent specimen outputs.

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