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EXHIBIT 112

Edmontonia

Edmontonia

Edmontonia was a Late Cretaceous North American plant-eater protected by bony plates and lateral spikes. [1] [4]

Evidence profile
PERIODLate Cretaceous[2]
DIETHerbivoreEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDAlberta, Canada, Montana, USA[2]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Edmontonia

Edmontonia was a plant-eating armored dinosaur from Late Cretaceous North America. It belonged to the nodosaurs, part of the wider armored dinosaur group. A museum reference of about four metres offers an educational introduction without identifying a measured adult or a size range for every species. Its fossil record includes evidence about growth as well as appearance. [D1]

An early study of a Montana fossil describes neck armor and lateral spikes. Its incomplete tail material does not show a heavy club like that of some other armored dinosaurs. [D4]

Thin sections of another specimen's limb bones reveal extensive rebuilding of older tissue and relatively slow deposition near the outside. These microscopic features help researchers investigate advanced growth. They preserve a record of bone formation rather than a direct view of the animal's activities. [D5]

That growth evidence does not establish an exact age at breeding. It also cannot automatically identify the maturity of a different specimen used in a body-size model. Edmontonia demonstrates why evidence about one individual's bones must stay separate from claims about every adult in a genus. [D5]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Preserved neck armor shows how densely protected its front end could be. Its small teeth, limb bones and microscopic bone structure reveal different parts of its biology. [D4] [D5]

Profile scope: Genus profile covering E. longiceps and E. rugosidens; their shoulder-spike proportions differ. [D2]

Time
Late Cretaceous
Fossil locations in this exhibit
Alberta, Canada, Montana, USA
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Length

Inferred

About 4 m (museum reference)

This educational genus estimate has no named individual, reconstruction method or maturity assessment. It does not define both species or an adult size range. [D1]

Mass

Inferred

About 3.1 tonnes (composite model)

Paul models AMNH 5665 with USNM 11868 using volume and assumed density. Maturity is unspecified; this reconstruction combines material and is not a species-wide range. [D3]

Diet and feeding

Source supported

Plant-eater

The museum identifies herbivory; preserved small, ridged teeth are consistent with plant processing. Specific food plants remain unresolved. [D1] [D4]

Habitat and range

Source supported

Late Cretaceous Alberta and Montana

Reviewed fossils come from the Horseshoe Canyon, Dinosaur Park and Two Medicine formations, spanning Campanian to early Maastrichtian deposits. [D2]

Movement

Inferred

Four-legged walking inferred

The limb and trunk anatomy support a low quadrupedal reconstruction. No measured running speed is established. [D4]

Distinctive anatomy

Source supported

Neck armor and lateral spikes; unclubbed tail

E. rugosidens has larger lateral spikes than E. longiceps. These species differences matter when restoring the animal. [D2] [D4]

Reproduction & life together

Reproduction and care

Not established

Not established here

Limb histology records growth, not eggs, nests or parental care. Focused review established no precise reproductive age or breeding pattern. [D5]

Courtship and mating

Not established

Not established here

The armor does not by itself establish sex differences, mate selection or a courtship display. No genus-specific mating behavior was demonstrated in the reviewed sources.

Social behavior

Not established

Not established here

Reviewed associated skeletons establish individual anatomy, without demonstrating persistent herds, family groups or solitary behavior.

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. EdmontoniaNatural History Museum, London · Accessed 2026-10-02Museum profile inspected: 4 m educational length, herbivory, Late Cretaceous North American occurrence and nodosaur identification; individual and method unspecified. · Inspected: page
  2. Redescription of the Panoplosaurus mirus (Ankylosauria: Nodosauridae) holotype skull using computed tomography, and a new comparative scheme for panoplosaurin cranial ornamentationJournal of Vertebrate Paleontology; Livius et al. (2026) · Accessed 2026-10-02Published full 28-page PDF inspected, including comparative diagnoses and synonymies. Retains E. longiceps and E. rugosidens; distinguishes their lateral spines and records Canadian/Montana provenance. TMP 1998.098.0001 is Edmontonia sp., and ROM 1215 is Panoplosaurus. · Inspected: page
  3. The Princeton Field Guide to Dinosaurs, Second Edition — Mass TablePrinceton University Press; Gregory S. Paul · Accessed 2026-10-02Original 21-page model appendix inspected. Edmontonia combines AMNH 5665 and USNM 11868 for 3,070 kg using volume and assumed density. Adjacent ~7.75 is a femur/model scaling input in decimeters, not total body length; its unit heading misprints decameters. No explicit revision date. · Inspected: page
  4. On dinosaurian reptiles from the Two Medicine Formation of MontanaProceedings of the United States National Museum; Gilmore (1930) · Accessed 2026-10-02Original full article and plates inspected. USNM 11868 was named Palaeoscincus rugosidens, now E. rugosidens; neck armor, lateral spikes, teeth and incomplete unclubbed tail material are described. · Inspected: page
  5. Long Bone Histology and Growth Patterns in Ankylosaurs: Implications for Life History and EvolutionPLOS ONE; Stein, Hayashi and Sander (2013) · Accessed 2026-10-02Primary full text inspected. Associated TMP 1998.98.1 limb histology supports advanced growth and relatively slow bone deposition; its E. rugosidens species referral is broadened to Edmontonia sp. in the 2026 review. No nesting or precise reproductive age is established. · Inspected: page

THE STORY IN THE FOSSILS

Meet Edmontonia

Gilmore’s description documents neck plates and large lateral spikes in the Montana fossil now called E. rugosidens. Its preserved tail material does not carry the heavy club associated with some other armored dinosaurs. [4]

Microscopic sections from TMP 1998.98.1 show extensively remodeled limb bones and relatively slow deposition near the exterior. That growth evidence cannot supply an exact age at breeding or the maturity of a different size-model specimen. [5]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. EdmontoniaNatural History Museum, London · Accessed 2026-10-02Museum profile inspected: 4 m educational length, herbivory, Late Cretaceous North American occurrence and nodosaur identification; individual and method unspecified. · Inspected: page
  2. Redescription of the Panoplosaurus mirus (Ankylosauria: Nodosauridae) holotype skull using computed tomography, and a new comparative scheme for panoplosaurin cranial ornamentationJournal of Vertebrate Paleontology; Livius et al. (2026) · Accessed 2026-10-02Published full 28-page PDF inspected, including comparative diagnoses and synonymies. Retains E. longiceps and E. rugosidens; distinguishes their lateral spines and records Canadian/Montana provenance. TMP 1998.098.0001 is Edmontonia sp., and ROM 1215 is Panoplosaurus. · Inspected: page
  3. The Princeton Field Guide to Dinosaurs, Second Edition — Mass TablePrinceton University Press; Gregory S. Paul · Accessed 2026-10-02Original 21-page model appendix inspected. Edmontonia combines AMNH 5665 and USNM 11868 for 3,070 kg using volume and assumed density. Adjacent ~7.75 is a femur/model scaling input in decimeters, not total body length; its unit heading misprints decameters. No explicit revision date. · Inspected: page
  4. On dinosaurian reptiles from the Two Medicine Formation of MontanaProceedings of the United States National Museum; Gilmore (1930) · Accessed 2026-10-02Original full article and plates inspected. USNM 11868 was named Palaeoscincus rugosidens, now E. rugosidens; neck armor, lateral spikes, teeth and incomplete unclubbed tail material are described. · Inspected: page
  5. Long Bone Histology and Growth Patterns in Ankylosaurs: Implications for Life History and EvolutionPLOS ONE; Stein, Hayashi and Sander (2013) · Accessed 2026-10-02Primary full text inspected. Associated TMP 1998.98.1 limb histology supports advanced growth and relatively slow bone deposition; its E. rugosidens species referral is broadened to Edmontonia sp. in the 2026 review. No nesting or precise reproductive age is established. · Inspected: page

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