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EXHIBIT 177

Guanlong

Guanlong wucaii

Guanlong was a slender predator with long, three-fingered arms and a delicate crest along its skull. [4]

Evidence profile
PERIODLate Jurassic[1]
DIETCarnivoreEvidence and uncertainty in the dossier
TOTAL LENGTH3–3 mEstimate scope matters
FOSSILS FOUNDChina[1]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Guanlong

Guanlong was a Late Jurassic tyrannosauroid from China, an early relative of the group that later included Tyrannosaurus. It had relatively long arms and three clawed fingers on each hand, contrasting with the much smaller forelimbs of giant tyrannosaurids. Serrated teeth support a predatory interpretation. The exact prey and hunting method remain uncertain, rather than being established by its family name or a dramatic illustration. [D4]

Its median skull crest was exceptionally thin and contained air spaces. Much of the bony structure was only about 1.5 millimetres thick. Researchers proposed a display function, but the ornament alone does not establish sex differences or a courtship routine. [D1]

The fossils came from Wucaiwan in northwestern China, in a landscape interpreted as seasonal forest with rivers, conifers and ferns. Its species name refers to the colorful rocks there, not to preserved skin colors. Two nearly complete skeletons include an adult and a smaller animal. Their preservation offers unusually useful comparisons of growth, without revealing a lifetime of hunting or interactions. [D4]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Its two original skeletons preserve different growth stages. The crest’s display function is a hypothesis, and their burial association does not establish family life. [D1] [D2]

Profile scope: Genus — Guanlong; G. wucaii, with adult IVPP V14531 distinguished from growing V14532. [D1]

Time
Late Jurassic
Fossil locations in this exhibit
China
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Body length

Inferred

About 3 m (adult type reconstruction)

Original estimate for IVPP V14531; an individual reference rather than a species maximum. [D1]

Body mass

Inferred

About 72 kg (limb model)

IVPP V14531; measured femoral circumference, independently adult. [D5] [D1]

Diet and feeding

Inferred

Animal prey inferred

Serrated teeth support predatory feeding; particular prey, hunting style and stomach contents are not established. [D4]

Habitat and setting

Source supported

Upper Shishugou Formation, Xinjiang

Oxfordian deposits at Wucaiwan record a seasonal alluvial landscape with wet, locally trampled depressions. [D2]

Movement

Inferred

Two-legged; long grasping arms

Slender hindlimbs supported terrestrial movement. Three clawed fingers and relatively long arms contrast with the reduced forelimbs of giant tyrannosaurids. [D4]

Recognizable anatomy

Source supported

Thin, air-filled median crest

Much of the crest is only about 1.5 mm thick; pneumatic openings and slender supporting sheets form a fragile structure. [D1]

Reproduction & life together

Reproduction and nesting

Not established

Not established here

Histology distinguishes a twelve-year individual from a six-year growing one. These growth ages do not demonstrate first breeding, nests or parental care. [D1]

Mating and courtship

Inferred

Crest display proposed

Sexual selection was suggested for the delicate ornament. Sex differences and a mating display have not been demonstrated. [D1]

Social behavior

Not established

Not established here

Adult and younger skeletons occur in a mixed-species trap deposit. Sequential accumulation offers an explanation without requiring a family or hunting group. [D2]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. A basal tyrannosauroid dinosaur from the Late Jurassic of China (Xu et al., 2006)Nature · Accessed 2026-10-02Full original article and supporting information inspected in the IVPP-hosted 16-page PDF. Type IVPP V14531 is about 3 m long; fibular histology identifies a late stationary stage, dying in its twelfth year. V14532 died while growing in its sixth year. Oxfordian upper Shishugou, Wucaiwan, Xinjiang. Extremely thin pneumatic crest and three-fingered hand; sexual selection is proposed, not demonstrated. · Inspected: page
  2. Dinosaur death pits from the Jurassic of China (Eberth, Xu and Clark, 2010)PALAIOS; original author copy at IVPP · Accessed 2026-10-02Original sedimentology and taphonomic sections inspected. Sequential miring in wet, sauropod-trampled depressions is proposed for mixed dinosaur associations. Guanlong specimens at TBB 2002 do not establish the same social inference proposed for the separate, richer Limusaurus assemblage. · Inspected: page
  3. The skull of Monolophosaurus jiangi and its implications for early theropod evolution and phylogeny (Brusatte, Benson, Currie and Zhao, 2010)Zoological Journal of the Linnean Society; author manuscript at University of Edinburgh · Accessed 2026-10-02Full Guanlong–Monolophosaurus identity discussion inspected in author manuscript pp. 69–73. Explicitly rejects Carr’s historical immature-Monolophosaurus suggestion using adult histology and numerous distinguishing characters; Guanlong remains a distinct tyrannosauroid. · Inspected: page
  4. Dinosaur — Guanlong wucaii (Australian Museum)Australian Museum · Accessed 2026-10-02Institutional fact sheet inspected: long arms, three fingers, predatory interpretation, Late Jurassic China and seasonal forest setting. Its educational 4 m length differs from the original approximately 3 m type reconstruction; speculative colors and food relationships are not adopted. · Inspected: page
  5. Rates of dinosaur body mass evolution indicate 170 million years of sustained ecological innovation on the avian stem lineage (Benson et al., 2014), Dataset S1PLOS Biology · Accessed 2026-10-02Original XLS row inspected: adult IVPP V14531, measured femoral circumference 104 mm, femur length 408 mm, model mass 71.587864836172 kg. Adult identification is independently established by Xu et al. 2006 histology. · Inspected: page

THE STORY IN THE FOSSILS

Two growth stages in a fossil trap.

The burial interpretation matters: a younger skeleton below an older one can reflect separate accidents. It is insufficient evidence for parenting or pack behavior. [2]

The original adult growth diagnosis also helps reject the historical claim that Guanlong was merely a young Monolophosaurus. [3]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. A basal tyrannosauroid dinosaur from the Late Jurassic of China (Xu et al., 2006)Nature · Accessed 2026-10-02Full original article and supporting information inspected in the IVPP-hosted 16-page PDF. Type IVPP V14531 is about 3 m long; fibular histology identifies a late stationary stage, dying in its twelfth year. V14532 died while growing in its sixth year. Oxfordian upper Shishugou, Wucaiwan, Xinjiang. Extremely thin pneumatic crest and three-fingered hand; sexual selection is proposed, not demonstrated. · Inspected: page
  2. Dinosaur death pits from the Jurassic of China (Eberth, Xu and Clark, 2010)PALAIOS; original author copy at IVPP · Accessed 2026-10-02Original sedimentology and taphonomic sections inspected. Sequential miring in wet, sauropod-trampled depressions is proposed for mixed dinosaur associations. Guanlong specimens at TBB 2002 do not establish the same social inference proposed for the separate, richer Limusaurus assemblage. · Inspected: page
  3. The skull of Monolophosaurus jiangi and its implications for early theropod evolution and phylogeny (Brusatte, Benson, Currie and Zhao, 2010)Zoological Journal of the Linnean Society; author manuscript at University of Edinburgh · Accessed 2026-10-02Full Guanlong–Monolophosaurus identity discussion inspected in author manuscript pp. 69–73. Explicitly rejects Carr’s historical immature-Monolophosaurus suggestion using adult histology and numerous distinguishing characters; Guanlong remains a distinct tyrannosauroid. · Inspected: page
  4. Dinosaur — Guanlong wucaii (Australian Museum)Australian Museum · Accessed 2026-10-02Institutional fact sheet inspected: long arms, three fingers, predatory interpretation, Late Jurassic China and seasonal forest setting. Its educational 4 m length differs from the original approximately 3 m type reconstruction; speculative colors and food relationships are not adopted. · Inspected: page
  5. Rates of dinosaur body mass evolution indicate 170 million years of sustained ecological innovation on the avian stem lineage (Benson et al., 2014), Dataset S1PLOS Biology · Accessed 2026-10-02Original XLS row inspected: adult IVPP V14531, measured femoral circumference 104 mm, femur length 408 mm, model mass 71.587864836172 kg. Adult identification is independently established by Xu et al. 2006 histology. · Inspected: page

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