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EXHIBIT 198

Kulindadromeus

Kulindadromeus zabaikalicus

Kulindadromeus combined simple and compound feather-like appendages with scales on distinct regions of its small, two-legged body. [1]

Evidence profile
PERIODMiddle Jurassic[2]
DIETHerbivoreEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDRussia[2]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Kulindadromeus

Kulindadromeus was a small, two-legged dinosaur from Siberia with teeth adapted for eating plants. Its hind legs were longer than its forelimbs. The discovery became especially interesting because preserved body coverings accompanied the bones. This gave researchers an opportunity to examine the surface of an animal that belonged outside the familiar feathered theropod group. [S1]

The covering was a mixture rather than one uniform coat. Short bristles occurred on the head and body, more complex structures on the arms and legs, and scales on parts of the tail and lower legs. Different regions therefore carried different structures. The fossils support that patchwork, but an illustration's overall colors and the appearance of unpreserved regions still require interpretation. [S1]

Later geological work places the locality in the Middle Jurassic and describes a lake setting with conifer-rich surroundings. Pollen and zircon evidence help establish that regional age. A date from volcanic particles is not automatically the precise date of a dinosaur's death. The environmental evidence adds a landscape to the anatomical picture while keeping its own dating limits clear. [D2]

Overview sources
  1. Fossils found in Siberia suggest all dinosaurs had feathersUniversity of Bristol · Accessed 2026-10-04Inspected: page

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Its integument expands the evidence for projecting appendages beyond theropod dinosaurs. Preservation varies by body region; a complete uniform coat is not observed. [D1]

Profile scope: Genus — Kulindadromeus; K. zabaikalicus, type partial skull INREC K3/109 and fragmentary referred material. Competing Kulinda names are conservatively reserved. [D1] [D3]

Time
Middle Jurassic
Fossil locations in this exhibit
Russia
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Body length

Inferred

About 1.5 m (original reconstruction)

The naming paper reconstructs the small biped from fragmentary material; it does not establish a complete adult measurement or species maximum. [D1]

Body mass

Inferred

About 2 kg (composite volume model)

Paul’s INREC composite model uses volume and density, with limb scaling. It does not identify a single complete individual or establish maturity. [D7]

Diet and feeding

Inferred

Herbivory inferred

Plant-adapted teeth support vegetation feeding. Specific plant preferences and stomach contents are not established by the reviewed description. [D3]

Habitat and setting

Source supported

Bathonian lake deposits, Siberia

Basal Ukurey Formation, Olov Depression. Zircon ages constrain a maximum depositional age; palynology supports the Middle Jurassic Bathonian assignment. [D2]

Movement

Inferred

Terrestrial biped inferred

Long hindlimbs, short forelimbs and an elongate balancing tail support a two-legged reconstruction. No measured speed or genus-specific trackway is established here. [D3]

Recognizable anatomy

Source supported

Regional scales and feather-like appendages

Simple head/trunk filaments, compound upper-limb structures and scaly distal legs/tail occur in separate preserved patches. Neither uniform plumage nor color is established. [D1]

Reproduction & life together

Reproduction and nesting

Not established

Not established here

The type skull probably represents an immature animal. Growing bones do not establish breeding age; focused searches found no attributable eggs, nests or parent–young association. [D1]

Mating and courtship

Inferred

Visual display proposed

The chapter suggests display for organized ribbon-like appendages, alongside other possible functions. This does not demonstrate sex differences, mate choice or a courtship sequence. [D3]

Social behavior

Not established

Not established here

Young individuals dominate the bonebeds. Attritional accumulation is proposed but needs testing; this association does not directly establish a herd or family group. [D3]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. A Jurassic ornithischian dinosaur from Siberia with both feathers and scales (Godefroit et al., 2014)Science 345(6195):451–455; original author copy at Royal Belgian Institute of Natural Sciences · Accessed 2026-10-02Full original article and 28-page Bristol-hosted supplement inspected. Names Kulindadromeus from INREC K3/109 partial skull; probable immature skull features are described in the supplement. Approximately 1.5 m bipedal herbivore, mostly isolated/partly articulated bones and regionally preserved appendages. Original Middle–Late Jurassic age is superseded by 2019. Main original PDF and supplement are saved locally; no complete adult individual is implied. · Inspected: page
  2. The rise of feathered dinosaurs: Kulindadromeus zabaikalicus, the oldest dinosaur with “feather-like” structures (Cincotta et al., 2019)PeerJ 7:e6239 · Accessed 2026-10-02Original full 20-page article inspected via the institutional PDF. Kulinda basal Ukurey Formation, Olov Depression, Transbaikalia. Palynology and detrital zircons support Bathonian age; 172.8±1.6 Ma is a maximum volcaniclastic age, not a direct dinosaur date. Discusses conifer-rich lake setting. Authors propose the other Kulinda taxa are dubious/probable synonyms; their spelling Lepidocheirosaurus natalis differs from the actual 2015 natatilis and is not treated as a distinct species. · Inspected: page
  3. Integumentary Structures in Kulindadromeus zabaikalicus, a Basal Neornithischian Dinosaur from the Jurassic of Siberia (Godefroit et al., 2020)The Evolution of Feathers, Springer, pp 47–65; original institutional author copy · Accessed 2026-10-02Full 19-page chapter inspected, especially sections 4.2–4.6. Reviews skeletal and regionally preserved integumentary anatomy; simple filaments, compound tufts and ribbon-like structures. Visual display, balance and egg insulation are proposed functions, not observed behavior. Mostly young individuals motivate an attritional-accumulation hypothesis requiring age-distribution/taphonomic tests. Supports probable overlap of competing Kulinda names but acknowledges specimen association and taxonomic difficulties. · Inspected: page
  4. Two new ornithischian dinosaurs (Hypsilophodontia, Ornithopoda) from the Late Jurassic of Russia (Alifanov and Saveliev, 2014)Paleontological Journal 48:414–425 · Accessed 2026-10-02Original publisher abstract and issue metadata inspected, not full diagnoses. Names Kulindapteryx ukureica and Daurosaurus olovus from Kulinda and assigns them to different families. English publication 6 August 2014; the exact Russian-print date relative to 25 July Science naming was not established, so nomenclatural priority is not declared. · Inspected: abstract
  5. New Data on Ornithischian Dinosaurs from the Late Jurassic of Transbaikalia, Russia (Alifanov, 2018)Paleontological Journal 52:1633–1636 · Accessed 2026-10-02Original publisher abstract inspected. Explicitly contests the monospecific interpretation and diagnostic clarity of Kulindadromeus, retaining Daurosaurus and Kulindapteryx as described alternatives. This competing interpretation is recorded alongside 2019/2020 probable-synonym proposals, rather than erased. · Inspected: abstract
  6. The most ancient Ornithomimosaur (Theropoda, Dinosauria), with cover imprints from the Upper Jurassic of Russia (Alifanov and Saveliev, 2015)Paleontological Journal 49:636–650 · Accessed 2026-10-02Original publisher abstract inspected, not full osteology. Names Lepidocheirosaurus natatilis from the Kulinda assemblage and interprets it as an ornithomimosaur. That interpretation conflicts with the later tentative Kulindadromeus referral; the genus is reserved rather than marked a settled synonym. · Inspected: abstract
  7. Author mass-model appendix to The Princeton Field Guide to Dinosaurs (Paul; current publisher file)Princeton University Press · Accessed 2026-10-02Original 21-page publisher appendix inspected: page 1 defines model volume×specific gravity and limb-length scaling; page 8 lists “INREC composite:2:~1.00” under Kulindadromeus. Two is mass in kilograms; the last number is limb-bone length, not total body length. No individual identity or maturity is established. The header’s decameters unit is an apparent typo for decimeters; no limb conversion is used here. · Inspected: page

THE STORY IN THE FOSSILS

One fossil site, competing taxonomic interpretations.

The original competing descriptions named Daurosaurus and Kulindapteryx. Their later proposed overlap requires an explicit reservation, rather than two additional exhibits. [4] [2]

A later author disputed the assumption of one Kulinda dinosaur taxon. That published disagreement limits how confidently disarticulated remains can be assembled into one animal. [5]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. A Jurassic ornithischian dinosaur from Siberia with both feathers and scales (Godefroit et al., 2014)Science 345(6195):451–455; original author copy at Royal Belgian Institute of Natural Sciences · Accessed 2026-10-02Full original article and 28-page Bristol-hosted supplement inspected. Names Kulindadromeus from INREC K3/109 partial skull; probable immature skull features are described in the supplement. Approximately 1.5 m bipedal herbivore, mostly isolated/partly articulated bones and regionally preserved appendages. Original Middle–Late Jurassic age is superseded by 2019. Main original PDF and supplement are saved locally; no complete adult individual is implied. · Inspected: page
  2. The rise of feathered dinosaurs: Kulindadromeus zabaikalicus, the oldest dinosaur with “feather-like” structures (Cincotta et al., 2019)PeerJ 7:e6239 · Accessed 2026-10-02Original full 20-page article inspected via the institutional PDF. Kulinda basal Ukurey Formation, Olov Depression, Transbaikalia. Palynology and detrital zircons support Bathonian age; 172.8±1.6 Ma is a maximum volcaniclastic age, not a direct dinosaur date. Discusses conifer-rich lake setting. Authors propose the other Kulinda taxa are dubious/probable synonyms; their spelling Lepidocheirosaurus natalis differs from the actual 2015 natatilis and is not treated as a distinct species. · Inspected: page
  3. Integumentary Structures in Kulindadromeus zabaikalicus, a Basal Neornithischian Dinosaur from the Jurassic of Siberia (Godefroit et al., 2020)The Evolution of Feathers, Springer, pp 47–65; original institutional author copy · Accessed 2026-10-02Full 19-page chapter inspected, especially sections 4.2–4.6. Reviews skeletal and regionally preserved integumentary anatomy; simple filaments, compound tufts and ribbon-like structures. Visual display, balance and egg insulation are proposed functions, not observed behavior. Mostly young individuals motivate an attritional-accumulation hypothesis requiring age-distribution/taphonomic tests. Supports probable overlap of competing Kulinda names but acknowledges specimen association and taxonomic difficulties. · Inspected: page
  4. Two new ornithischian dinosaurs (Hypsilophodontia, Ornithopoda) from the Late Jurassic of Russia (Alifanov and Saveliev, 2014)Paleontological Journal 48:414–425 · Accessed 2026-10-02Original publisher abstract and issue metadata inspected, not full diagnoses. Names Kulindapteryx ukureica and Daurosaurus olovus from Kulinda and assigns them to different families. English publication 6 August 2014; the exact Russian-print date relative to 25 July Science naming was not established, so nomenclatural priority is not declared. · Inspected: abstract
  5. New Data on Ornithischian Dinosaurs from the Late Jurassic of Transbaikalia, Russia (Alifanov, 2018)Paleontological Journal 52:1633–1636 · Accessed 2026-10-02Original publisher abstract inspected. Explicitly contests the monospecific interpretation and diagnostic clarity of Kulindadromeus, retaining Daurosaurus and Kulindapteryx as described alternatives. This competing interpretation is recorded alongside 2019/2020 probable-synonym proposals, rather than erased. · Inspected: abstract
  6. The most ancient Ornithomimosaur (Theropoda, Dinosauria), with cover imprints from the Upper Jurassic of Russia (Alifanov and Saveliev, 2015)Paleontological Journal 49:636–650 · Accessed 2026-10-02Original publisher abstract inspected, not full osteology. Names Lepidocheirosaurus natatilis from the Kulinda assemblage and interprets it as an ornithomimosaur. That interpretation conflicts with the later tentative Kulindadromeus referral; the genus is reserved rather than marked a settled synonym. · Inspected: abstract
  7. Author mass-model appendix to The Princeton Field Guide to Dinosaurs (Paul; current publisher file)Princeton University Press · Accessed 2026-10-02Original 21-page publisher appendix inspected: page 1 defines model volume×specific gravity and limb-length scaling; page 8 lists “INREC composite:2:~1.00” under Kulindadromeus. Two is mass in kilograms; the last number is limb-bone length, not total body length. No individual identity or maturity is established. The header’s decameters unit is an apparent typo for decimeters; no limb conversion is used here. · Inspected: page

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