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EXHIBIT 201

Meganeura

Meganeura

Prehistoric insect

Meganeura was a large flying insect from Carboniferous France, known for its expansive wings. [1]

Evidence profile
PERIODLate Carboniferous[1]
DIETCarnivoreEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDFrance[1]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Meganeura

Meganeura was an insect that flew over what is now France late in the Carboniferous Period, roughly 300 million years ago. The best-known species, Meganeura monyi, came from the coalfields of Commentry. Its wings could spread about 70 centimeters across. That number describes the distance between the wing tips, rather than the length of its body. It was a large insect, not a dinosaur. [D1]

Its fossils belong to a much larger collection of ancient insects. Workers and researchers recovered many specimens from the old Commentry coal workings, and museums kept them for later study. An insect wing can preserve a fine network of veins even when the rest of the body is missing. Those patterns help researchers recognize kinds of insects and reconsider names assigned long ago. [D2]

Spines on the legs fit a predatory lifestyle, but the clearest detailed evidence comes from a related insect called Meganeurites. Comparing relatives helps build a careful picture without making every feature certain. The reviewed fossils do not tell us how Meganeura courted a mate or exactly where it laid eggs. [D4]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Commentry collecting produced a remarkable insect archive; isolated wings make later identification and revision especially important. [D2]

Profile scope: Genus — Meganeura; reconstruction anchored to M. monyi from Commentry, France. [D1]

Time
Late Carboniferous
Fossil locations in this exhibit
France
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Size reference

Source supported

About 70 cm wingspan

Museum reference for M. monyi; wing spread is not body length or a genus-wide adult range. [D1]

Body mass

Not established

Not established here

The inspected museum account supplies no defensible fossil-individual body-mass estimate. [D1]

Diet

Inferred

Predatory feeding inferred

Spiny legs and comparison with better-preserved meganeurids support carnivory; the detailed mouthpart study concerns Meganeurites. [D4]

Habitat

Inferred

Commentry coal-basin surroundings

Fossils come from a limnic basin; open woodland margins are a comparative habitat interpretation, not a preserved home. [D4]

Movement

Inferred

Wing-powered flight

Large wings support flight; precise speed and modern dragonfly maneuvers are not established. [D4]

Distinctive anatomy

Source supported

Veined wings and spiny legs

Head details are poorly preserved; much of the complete living outline needs comparison with related insects. [D4]

Reproduction & life together

Reproduction

Not established

Not established here

The reviewed anatomical study does not identify attributable eggs or a Meganeura life-cycle series. [D4]

Courtship

Not established

Not established here

No genus-specific courtship sequence is established by the reviewed fossils. [D4]

Social behavior

Not established

Not established here

Shared burial does not demonstrate coordinated groups or social organization. [D2]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. Meganeura monyi, libellule géanteMuséum national d’Histoire naturelle · Accessed 2026-10-05Museum specimen profile inspected: Commentry occurrence, Late Carboniferous age, discovery and approximately 70 cm wingspan. Its absolute largest-insect wording is not adopted because Meganeuropsis is larger. · Inspected: page
  2. Fossil insect collectionMuséum national d’Histoire naturelle · Accessed 2026-10-05Museum collection account inspected: historical Commentry collecting, Meganeura monyi, isolated wings and later taxonomic revisions. · Inspected: page
  3. MNHN.F.R51142 — historical Meganeura monyi determinationsMuséum national d’Histoire naturelle specimen database · Accessed 2026-10-05Full specimen record and determination history inspected. Header retains Dictyoneura monyi; later history identifies Meganeura monyi. Database calls the incomplete specimen a lectotype, while later papers use holotype; the exhibit avoids choosing between these conflicting type labels. · Inspected: page
  4. Palaeozoic giant dragonflies were hawker predators (Nel et al., 2018)Scientific Reports 8:12141 · Accessed 2026-10-05Full article inspected, particularly morphology and ecological interpretation. Direct new anatomical evidence is Meganeurites gracilipes, not Meganeura or Meganeuropsis; predator and open-space comparisons for those genera are explicitly inferences. Reviewed Meganeura head preservation and known leg spines. · Inspected: page

THE STORY IN THE FOSSILS

A wing from the coalfields

The Commentry collection shows how an insect can become familiar through wings while much of its living biology remains difficult to reconstruct. [2]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. Meganeura monyi, libellule géanteMuséum national d’Histoire naturelle · Accessed 2026-10-05Museum specimen profile inspected: Commentry occurrence, Late Carboniferous age, discovery and approximately 70 cm wingspan. Its absolute largest-insect wording is not adopted because Meganeuropsis is larger. · Inspected: page
  2. Fossil insect collectionMuséum national d’Histoire naturelle · Accessed 2026-10-05Museum collection account inspected: historical Commentry collecting, Meganeura monyi, isolated wings and later taxonomic revisions. · Inspected: page
  3. MNHN.F.R51142 — historical Meganeura monyi determinationsMuséum national d’Histoire naturelle specimen database · Accessed 2026-10-05Full specimen record and determination history inspected. Header retains Dictyoneura monyi; later history identifies Meganeura monyi. Database calls the incomplete specimen a lectotype, while later papers use holotype; the exhibit avoids choosing between these conflicting type labels. · Inspected: page
  4. Palaeozoic giant dragonflies were hawker predators (Nel et al., 2018)Scientific Reports 8:12141 · Accessed 2026-10-05Full article inspected, particularly morphology and ecological interpretation. Direct new anatomical evidence is Meganeurites gracilipes, not Meganeura or Meganeuropsis; predator and open-space comparisons for those genera are explicitly inferences. Reviewed Meganeura head preservation and known leg spines. · Inspected: page

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