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EXHIBIT 110

Shantungosaurus

Shantungosaurus giganteus

Shantungosaurus was a giant plant-eating hadrosaur from China, with a broad beak and deep dental batteries rather than an elaborate hollow crest. [2] [1]

Evidence profile
PERIODLate Cretaceous[1]
DIETHerbivoreEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDChina[1]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Shantungosaurus

Shantungosaurus was a large plant-eating dinosaur from Late Cretaceous China. A museum reference of about fifteen metres conveys an impressive scale without identifying a complete measured adult. Different reconstructions can use different body outlines, so a headline length should not automatically be paired with a weight estimate calculated for a different outline. [D2]

At Kugou, a remarkable accumulation of disarticulated bones includes 110 thigh bones, implying at least 55 individuals. Their similar sizes and fused skeletal features support an interpretation of an adult aggregation. Rapid burial and the jumbled remains still limit the picture. An adult group gathering at one place is not the same as proof of a permanent herd. [D4]

The wider region also preserves dinosaur eggs, but egg discoveries and named body fossils are documented separately. Those eggs cannot securely be assigned to Shantungosaurus just because they came from the same general area. The distinction keeps reproductive behavior honest: an ancient environment can contain many dinosaurs, and identifying which animal laid a particular egg requires evidence beyond proximity. [D5]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Cranial comparisons link it closely to Edmontosaurus while retaining a distinct genus. The reviewed material includes multiple individuals, and proposed adult grouping requires a taphonomic interpretation. [D1] [D4]

Profile scope: Genus profile centered on Shantungosaurus giganteus from Shandong. Zhuchengosaurus and Huaxiaosaurus are included as reviewed junior synonyms; composite mounts and outline models are not individual adult ranges. [D1]

Time
Late Cretaceous
Fossil locations in this exhibit
China
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Body length

Inferred

About 15 m; museum reference

The directory provides no individual or maturity assessment. A historical mass model uses a separate 17 m outline; neither is a directly measured adult species maximum. [D2]

Body mass

Inferred

About 22.5 tonnes; historical outline model

Seebacher’s model integrates a reconstructed 17 m outline using width and density assumptions. No inventory or maturity is specified; it is not the mass of the museum’s 15 m reference. [D3]

Diet

Inferred

Plants

The museum identifies herbivory. Extensive dental batteries processed vegetation, but no specific diet of named plants or daily intake is established here. [D2] [D1]

Habitat & fossil setting

Source supported

Wangshi Group, Shandong

The systematic revision places material from the upper Xingezhuang to middle Hongtuya formations in the middle–late Campanian. This supports Late Cretaceous China; individual habitat preferences remain unresolved. [D1]

Movement

Inferred

Quadrupedal walking plausible; bipedality possible

Hadrosaur limb anatomy supports four-legged walking at group level. Hone’s author account sees no reason to exclude bipedal movement; neither source measures Shantungosaurus gait frequency or speed. [D6] [D4]

Distinctive anatomy

Source supported

Broad beak and deep dental batteries

The revision records up to seven teeth per alveolus in the middle lower-jaw row, including replacement teeth. This is not seven simultaneously functional teeth at every position. [D1]

Reproduction & life together

Reproduction & nesting

Not established

Not established here

The regional review lists dinosaur eggs separately from body-fossil taxa. It does not establish a Shantungosaurus clutch, nest design, incubation strategy or parental care. [D5]

Courtship & mating

Not established

Not established here

Reviewed skull anatomy and focused searches do not establish mating displays, calls, mate choice or sexual dimorphism. Crest absence alone cannot answer these questions.

Social behavior

Inferred

Adult aggregation inferred

Hone reports 110 femora, implying at least 55 individuals at Kugou. Narrow size distribution and fused sacra support an adult-group interpretation; rapid burial does not prove permanent herds or juvenile care. [D4]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. Comparative Osteology and Phylogenetic Relationship of Edmontosaurus and Shantungosaurus from the Upper Cretaceous of North America and East AsiaActa Geologica Sinica; Xing and colleagues (2014), author-deposited published paper · Accessed 2026-10-02Full-paper excerpts: Wangshi Group distribution, middle–late Campanian chronology, skull/dental characters and synonymy of Zhuchengosaurus maximus and Huaxiaosaurus aigahtens. · Inspected: excerpt
  2. ShantungosaurusNatural History Museum, London · Accessed 2026-10-02Late Cretaceous China, herbivory and 15 m museum reference; no individual, maturity or reconstruction method specified. · Inspected: page
  3. A new method to calculate allometric length-mass relationships of dinosaursJournal of Vertebrate Paleontology; Seebacher (2001), author-deposited full text · Accessed 2026-10-02Methods and Table 2 inspected: reconstructed 17 m outline and 22467.1 kg, without an individual inventory or maturity assessment. · Inspected: excerpt
  4. A population of Shantungosaurus, the largest ornithischianDave Hone (2014); primary author’s account of his Hadrosaurs research chapter · Accessed 2026-10-02Author’s full research account inspected, not the inaccessible book chapter: Kugou femur counts, adult interpretation, rapid burial and proposed aggregation. · Inspected: page
  5. Review of historical and current research on the Late Cretaceous dinosaurs and dinosaur eggs from Laiyang, ShandongVertebrata PalAsiatica; Zhang and colleagues (2017), IVPP-hosted published paper · Accessed 2026-10-02Body-fossil and egg-fauna tables and regional research history inspected. Regional eggs are classified separately and are not securely assigned to Shantungosaurus in this review. · Inspected: excerpt
  6. The evolution of quadrupedality in ornithischian dinosaursJournal of Iberian Geology; Maidment & Barrett (2017) · Accessed 2026-10-02Hadrosaur limb and quadrupedality discussion: broader-group functional inference, not a genus-specific measured gait or speed. · Inspected: excerpt

THE STORY IN THE FOSSILS

Meet Shantungosaurus.

Shantungosaurus is distinct from Edmontosaurus, although their skulls and tooth batteries share detailed features. Treating Zhuchengosaurus and Huaxiaosaurus as reviewed synonyms avoids counting the same genus three times. [1]

Kugou preserves thousands of disarticulated bones. Their size distribution suggests a remarkable adult aggregation, while the jumble of remains and burial process limit how much of everyday social life can be reconstructed. [4]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. Comparative Osteology and Phylogenetic Relationship of Edmontosaurus and Shantungosaurus from the Upper Cretaceous of North America and East AsiaActa Geologica Sinica; Xing and colleagues (2014), author-deposited published paper · Accessed 2026-10-02Full-paper excerpts: Wangshi Group distribution, middle–late Campanian chronology, skull/dental characters and synonymy of Zhuchengosaurus maximus and Huaxiaosaurus aigahtens. · Inspected: excerpt
  2. ShantungosaurusNatural History Museum, London · Accessed 2026-10-02Late Cretaceous China, herbivory and 15 m museum reference; no individual, maturity or reconstruction method specified. · Inspected: page
  3. A new method to calculate allometric length-mass relationships of dinosaursJournal of Vertebrate Paleontology; Seebacher (2001), author-deposited full text · Accessed 2026-10-02Methods and Table 2 inspected: reconstructed 17 m outline and 22467.1 kg, without an individual inventory or maturity assessment. · Inspected: excerpt
  4. A population of Shantungosaurus, the largest ornithischianDave Hone (2014); primary author’s account of his Hadrosaurs research chapter · Accessed 2026-10-02Author’s full research account inspected, not the inaccessible book chapter: Kugou femur counts, adult interpretation, rapid burial and proposed aggregation. · Inspected: page
  5. Review of historical and current research on the Late Cretaceous dinosaurs and dinosaur eggs from Laiyang, ShandongVertebrata PalAsiatica; Zhang and colleagues (2017), IVPP-hosted published paper · Accessed 2026-10-02Body-fossil and egg-fauna tables and regional research history inspected. Regional eggs are classified separately and are not securely assigned to Shantungosaurus in this review. · Inspected: excerpt
  6. The evolution of quadrupedality in ornithischian dinosaursJournal of Iberian Geology; Maidment & Barrett (2017) · Accessed 2026-10-02Hadrosaur limb and quadrupedality discussion: broader-group functional inference, not a genus-specific measured gait or speed. · Inspected: excerpt

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