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EXHIBIT 197

Tethyshadros

Tethyshadros insularis

Tethyshadros was a plant-eating hadrosauroid with dental batteries, an unusual tail and three tightly grouped hand digits. [4]

Evidence profile
PERIODLate Cretaceous[2]
DIETHerbivoreEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDItaly[2]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Tethyshadros

Tethyshadros is known from unusually complete, articulated dinosaur material from northeastern Italy. Articulated means many bones remained arranged together, helping researchers reconstruct how the skeleton fitted. It belongs close to the familiar duck-billed dinosaurs but outside the more narrowly defined family Hadrosauridae. That distinction matters: an animal can resemble members of a family without being placed inside it by a scientific analysis. [D1]

Its jaws contained many teeth arranged in stacked rows, a dental battery suited to processing plant food. The hand had three fingers and relatively limited movement. These preserved details help explain its feeding and body structure. They do not identify a favorite plant, reveal the sex of the fossil individual or tell us how the animals behaved together. [D4]

Research on its limbs and hands supports the ability to use both two-legged and four-legged postures. It does not establish an exact running speed. Whole-body restorations also depend on how missing or incomplete parts, especially the tail, are reconstructed. A measured preserved skeleton and a model's estimated complete length answer different questions, and reconstruction estimates retain those limits. [D3]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

Rib histology identifies “Antonio” as subadult and “Bruno” as older. Continued slow growth in Bruno remains possible. [D2]

Profile scope: Genus — Tethyshadros; T. insularis from northeastern Italy, a hadrosauroid outside Hadrosauridae. [D1]

Time
Late Cretaceous
Fossil locations in this exhibit
Italy
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Body length

Inferred

3.62 m preserved (subadult type)

SC57021 lacks the tail tip. A historical 4.51 m restoration is challenged by 2021’s shorter caudal series; neither is an adult maximum. [D3] [D2]

Body mass

Inferred

About 514–584 kg (older-individual model)

SC57247: isometric scaling from the type’s limb model using 15–20% larger linear dimensions. Continued slow growth remains possible; this is not a species range. [D2]

Diet and feeding

Inferred

Plant feeding inferred

Dental batteries held many teeth in vertical rows. This supports herbivory without identifying particular plants or an observed meal. [D4]

Habitat and setting

Source supported

Campanian carbonate-platform deposits

Villaggio del Pescatore, northeastern Italy; approximately 81.5–80.5 Ma. Marginal-marine burial deposits do not establish an aquatic lifestyle. [D2]

Movement

Inferred

Both two- and four-legged capability inferred

The long tibia and pillar-like hand support mixed capabilities. Digits II/III have hoof-like unguals; IV lacks one. Exclusive gait and speed remain unestablished. [D3]

Recognizable anatomy

Source supported

Three hand digits; specialized dental batteries

Manual digits I and V are absent. II–IV form a hand with limited mobility; complete tail length remains reconstructed. [D4]

Reproduction & life together

Reproduction and nesting

Inferred

Growth and clutch hypotheses

Slower bone growth suggests sexual maturation. Tail-based longer-oviduct/larger-clutch proposals remain untested by eggs, nests or reproductive tissues. [D2] [D3]

Mating and courtship

Not established

Not established here

A posterior first chevron inspired historical sex assignments, but the museum explicitly finds these unconfirmed. No inspected display sequence or mating behavior is documented. [D4]

Social behavior

Not established

Not established here

Several individuals occur at the fossil site. Their occurrence does not itself establish a herd, family or cooperative behavior; focused searches found no direct social test. [D2]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. Tethyshadros insularis, a new hadrosauroid dinosaur (Ornithischia) from the Upper Cretaceous of Italy (Dalla Vecchia, 2009)Journal of Vertebrate Paleontology · Accessed 2026-10-02Original publisher-indexed abstract inspected, not the subscription osteology. Names T. insularis from an articulated skeleton and places it near, but outside, Hadrosauridae. Its historical late Campanian–early Maastrichtian dating and island-dwarf interpretation are superseded or challenged by the inspected 2021 revision. · Inspected: abstract
  2. An Italian dinosaur Lagerstätte reveals the tempo and mode of hadrosauriform body size evolution (Chiarenza et al., 2021)Scientific Reports 11:23295 · Accessed 2026-10-02Original full article, histology, revised diagnosis, stratigraphy and downloaded DOCX supplement inspected. SC57021 Antonio is subadult; SC57247 Bruno is older with a tentative external fundamental system, but cortical drift/slow continued growth remain possible. Supplement S3 scales historical type mass 338.1776 kg by 1.15–1.20 cubed to give 514.33–584.37 kg, not a direct limb model of Bruno, whose humerus is missing. Bone-growth spacing supports a sexual-maturity hypothesis, not a measured breeding age. Supplement S2 rejects the historical 77-caudal model and tests approximately 44–51 elements. Several specimen-number typos in the original prose are not copied into this profile. · Inspected: page
  3. The unusual tail of Tethyshadros insularis (Dinosauria, Hadrosauroidea) from the Adriatic Island of the European Archipelago (Dalla Vecchia, 2020)Rivista Italiana di Paleontologia e Stratigrafia 126(3):583–628 · Accessed 2026-10-02Original 46-page article and eight-page supplement inspected, especially measurement, hand function, locomotion, reproductive hypotheses and conclusions. Type preserved length 3620 mm; historical 77-caudal restoration gives 4505.78 mm but is explicitly challenged by 2021. Manual II–IV are appressed with limited motion; II/III have hoof-like unguals, IV lacks one, I/V are absent. Locomotor traits support both postures without measured speed. Posterior first chevron/vent position motivates a longer-cloaca/oviduct and larger-clutch hypothesis; no reproductive tissues, eggs or clutch are directly demonstrated. · Inspected: page
  4. Dinosauri Antonio e Bruno (Museo Civico di Storia Naturale di Trieste)Museo Civico di Storia Naturale di Trieste · Accessed 2026-10-02Full institutional page inspected in Italian: dental batteries support herbivory, three manual digits, reduced hand mobility and uncertainty over Antonio’s sex. Its older adulthood, approximately 70 Ma, island-dwarf and 500 kg educational claims are not adopted against the 2021 revision. · Inspected: page

THE STORY IN THE FOSSILS

A growth-stage revision changes the interpretation.

Earlier skeletal fusion suggested adulthood, but rib histology revised Antonio to subadult. Bruno’s older status remains distinct from certainty that growth had ceased. [2]

The unusual hand is a support structure, rather than an assumed grasping hand. Its three grouped digits help distinguish the reconstruction from thumb-spiked iguanodontians. [3]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. Tethyshadros insularis, a new hadrosauroid dinosaur (Ornithischia) from the Upper Cretaceous of Italy (Dalla Vecchia, 2009)Journal of Vertebrate Paleontology · Accessed 2026-10-02Original publisher-indexed abstract inspected, not the subscription osteology. Names T. insularis from an articulated skeleton and places it near, but outside, Hadrosauridae. Its historical late Campanian–early Maastrichtian dating and island-dwarf interpretation are superseded or challenged by the inspected 2021 revision. · Inspected: abstract
  2. An Italian dinosaur Lagerstätte reveals the tempo and mode of hadrosauriform body size evolution (Chiarenza et al., 2021)Scientific Reports 11:23295 · Accessed 2026-10-02Original full article, histology, revised diagnosis, stratigraphy and downloaded DOCX supplement inspected. SC57021 Antonio is subadult; SC57247 Bruno is older with a tentative external fundamental system, but cortical drift/slow continued growth remain possible. Supplement S3 scales historical type mass 338.1776 kg by 1.15–1.20 cubed to give 514.33–584.37 kg, not a direct limb model of Bruno, whose humerus is missing. Bone-growth spacing supports a sexual-maturity hypothesis, not a measured breeding age. Supplement S2 rejects the historical 77-caudal model and tests approximately 44–51 elements. Several specimen-number typos in the original prose are not copied into this profile. · Inspected: page
  3. The unusual tail of Tethyshadros insularis (Dinosauria, Hadrosauroidea) from the Adriatic Island of the European Archipelago (Dalla Vecchia, 2020)Rivista Italiana di Paleontologia e Stratigrafia 126(3):583–628 · Accessed 2026-10-02Original 46-page article and eight-page supplement inspected, especially measurement, hand function, locomotion, reproductive hypotheses and conclusions. Type preserved length 3620 mm; historical 77-caudal restoration gives 4505.78 mm but is explicitly challenged by 2021. Manual II–IV are appressed with limited motion; II/III have hoof-like unguals, IV lacks one, I/V are absent. Locomotor traits support both postures without measured speed. Posterior first chevron/vent position motivates a longer-cloaca/oviduct and larger-clutch hypothesis; no reproductive tissues, eggs or clutch are directly demonstrated. · Inspected: page
  4. Dinosauri Antonio e Bruno (Museo Civico di Storia Naturale di Trieste)Museo Civico di Storia Naturale di Trieste · Accessed 2026-10-02Full institutional page inspected in Italian: dental batteries support herbivory, three manual digits, reduced hand mobility and uncertainty over Antonio’s sex. Its older adulthood, approximately 70 Ma, island-dwarf and 500 kg educational claims are not adopted against the 2021 revision. · Inspected: page

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