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EXHIBIT 181

Timurlengia

Timurlengia euotica

Timurlengia’s braincase reveals advanced tyrannosauroid sensory anatomy before the group reached giant body sizes. [4]

Evidence profile
PERIODLate Cretaceous[1]
DIETCarnivoreEvidence and uncertainty in the dossier
TOTAL LENGTHSee the dossierEstimate scope matters
FOSSILS FOUNDUzbekistan[1]
CompareRead the dossierView the sources

MEET THE ANIMAL

Meet Timurlengia

Timurlengia was a meat-eating relative of the tyrannosaurs, but it was much smaller than the giants that later made this family famous. Its fossils come from Uzbekistan. Sharp, blade-shaped teeth suggest that it sliced meat. A restored long-legged body is a useful picture of the animal, rather than one complete skeleton that was found together. [D3]

An important fossil is the braincase: the bones that surrounded the brain. Scans also allowed researchers to reconstruct parts of the inner ear. Its shape suggests sensitivity to low-frequency sounds. This helps explain how some advanced senses developed in tyrannosaur relatives before their bodies became enormous. It does not tell us what Timurlengia's calls sounded like. [D4]

Much of the rest of the picture comes from separate bones. Earlier studies described isolated jaws and other remains from different individuals, rather than a single animal with its whole body preserved. That distinction matters when imagining its appearance or daily life. Several fossils from an area do not, by themselves, show that these dinosaurs lived or hunted in a pack. [D2]

PREHISTORIC ANIMAL DOSSIER

A life in profile.

The animal, the evidence, and the questions still open.

A reconstructed inner ear provides evidence about hearing potential. Its missing body and provisional bone referrals prevent a measured adult whole-body profile. [D4] [D2]

Profile scope: Genus — Timurlengia; T. euotica, type braincase ZIN PH 1146/16 and separately referred isolated remains. [D1]

Time
Late Cretaceous
Fossil locations in this exhibit
Uzbekistan
How to read the evidence

Source supported identifies a description documented by the cited source. Inferred marks a reconstruction or functional estimate. Debated marks competing interpretations. Not established means the sources reviewed for this dossier do not support a firm conclusion; it does not mean a behavior never occurred.

A size reference is an estimate, not a population range. Nesting evidence does not by itself establish courtship, mate choice, or lifelong pairing.

Body length

Not established

Not established here

Only an isolated braincase and separately referred bones underpin the reconstruction. The inspected naming paper provides no measured whole-body length or adult maximum. [D1]

Body mass

Inferred

Approximately 170–270 kg (cranial comparison)

Braincase-size comparison with Xiongguanlong; the type may be immature. This is a provisional reconstruction, not an adult species range. [D1]

Diet and feeding

Inferred

Carnivory inferred

Slender, blade-like teeth were interpreted as suited to slicing meat. Suggested pursuit hunting and prey choices remain interpretations, without an observed meal. [D3]

Habitat and setting

Source supported

Bissekty Formation, Uzbekistan

Dzharakuduk in the Kyzylkum Desert; middle–upper Turonian, approximately 90–92 million years ago. [D1]

Movement

Inferred

Terrestrial biped inferred

Referred hindlimb material informs a provisional long-legged reconstruction. Speed is an author inference; separate bones do not certify one complete animal. [D2] [D3]

Recognizable anatomy

Inferred

Braincase and elongated cochlear duct

CT reconstruction supports sensitivity to low frequencies. Comparative research places advanced sensory anatomy before the evolution of giant tyrannosauroid bodies. [D4]

Reproduction & life together

Reproduction and nesting

Not established

Not established here

The braincase’s possible immaturity does not establish breeding age. Focused searches found no Timurlengia nest, eggs or reproductive tissues. [D1]

Mating and courtship

Not established

Not established here

An elongated inner-ear duct supports a hearing inference. It does not demonstrate mating calls, a display sequence or mate choice. [D4]

Social behavior

Not established

Not established here

The earlier skeletal sample contains isolated remains from separate individuals. Abundance across a formation does not itself establish a pack or family group. [D2]

Dossier research & sources

Sources and inspection notes for the claims in this profile.

  1. New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs (Brusatte et al., 2016)Proceedings of the National Academy of Sciences · Accessed 2026-10-02Original full material, diagnosis, CT and size discussion inspected. ZIN PH 1146/16 braincase from Dzharakuduk, Bissekty Formation, middle–upper Turonian, approximately 90–92 Ma. Size approximately 170–270 kg by comparison with Xiongguanlong; unfused/broken parabasisphenoid may indicate immaturity. Additional skull/postcranial bones are separate, not an associated whole skeleton. · Inspected: page
  2. Skeletal remains of Tyrannosauroidea from the Bissekty Formation at Dzharakuduk, Uzbekistan (Averianov and Sues, 2012)Cretaceous Research; original author copy at Smithsonian · Accessed 2026-10-02Original abstract, locality and skeletal description excerpts inspected from the 14-page institutional PDF. Isolated material was Tyrannosauroidea indeterminate before Timurlengia naming; juvenile/adult jaw fragments are distinct specimens. Taxonomic referral is supplied by the later 2016 paper, not assumed from the earlier title. · Inspected: page
  3. Newly found species reveals how T. rex became king of dinosaurs (University of Edinburgh, 2016)University of Edinburgh · Accessed 2026-10-02Original two-page institutional release fully inspected. Horse-sized reconstructed animal; sharp blade-like teeth, pursuit-hunting and speed are author interpretations. Mid-Cretaceous coastal plains are regional setting, not an observed hunt or habitat preference. · Inspected: page
  4. Neurosensory and Sinus Evolution as Tyrannosauroid Dinosaurs Developed Giant Size: Insights from the Endocranial Anatomy of Bistahieversor sealeyi (McKeown et al., 2020)The Anatomical Record · Accessed 2026-10-02Original full comparative sensory-evolution discussion inspected. Timurlengia is compared with larger tyrannosauroids: several sensory traits preceded gigantism. The focal specimen is Bistahieversor; its anatomy and sizes are not transferred to Timurlengia. · Inspected: page

THE STORY IN THE FOSSILS

A fossil ear before gigantic body size.

CT scans reconstruct internal sensory spaces that external skull shape alone cannot show. Their evolutionary comparison tests the timing of sensory changes, rather than the animal’s intelligence or social behavior. [4]

The original skeletal descriptions preceded naming Timurlengia. Keeping those isolated specimens separate prevents a composite reconstruction from becoming a fictitious complete fossil. [2]

A note on scale and time

The dossier identifies the scope of each reference estimate. A mount, an individual fossil and a species-wide adult range are different measurements. Comparison metrics remain unavailable where adult scope has not been established. Unknown does not mean zero.

Sources & editorial notes

Sources support the stated claims and scopes. Refer to the evidence notes for uncertainty.

  1. New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs (Brusatte et al., 2016)Proceedings of the National Academy of Sciences · Accessed 2026-10-02Original full material, diagnosis, CT and size discussion inspected. ZIN PH 1146/16 braincase from Dzharakuduk, Bissekty Formation, middle–upper Turonian, approximately 90–92 Ma. Size approximately 170–270 kg by comparison with Xiongguanlong; unfused/broken parabasisphenoid may indicate immaturity. Additional skull/postcranial bones are separate, not an associated whole skeleton. · Inspected: page
  2. Skeletal remains of Tyrannosauroidea from the Bissekty Formation at Dzharakuduk, Uzbekistan (Averianov and Sues, 2012)Cretaceous Research; original author copy at Smithsonian · Accessed 2026-10-02Original abstract, locality and skeletal description excerpts inspected from the 14-page institutional PDF. Isolated material was Tyrannosauroidea indeterminate before Timurlengia naming; juvenile/adult jaw fragments are distinct specimens. Taxonomic referral is supplied by the later 2016 paper, not assumed from the earlier title. · Inspected: page
  3. Newly found species reveals how T. rex became king of dinosaurs (University of Edinburgh, 2016)University of Edinburgh · Accessed 2026-10-02Original two-page institutional release fully inspected. Horse-sized reconstructed animal; sharp blade-like teeth, pursuit-hunting and speed are author interpretations. Mid-Cretaceous coastal plains are regional setting, not an observed hunt or habitat preference. · Inspected: page
  4. Neurosensory and Sinus Evolution as Tyrannosauroid Dinosaurs Developed Giant Size: Insights from the Endocranial Anatomy of Bistahieversor sealeyi (McKeown et al., 2020)The Anatomical Record · Accessed 2026-10-02Original full comparative sensory-evolution discussion inspected. Timurlengia is compared with larger tyrannosauroids: several sensory traits preceded gigantism. The focal specimen is Bistahieversor; its anatomy and sizes are not transferred to Timurlengia. · Inspected: page

FOLLOW YOUR CURIOSITY

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